Below is a list of my publications in order of release from oldest to newest; they include peer-reviewed articles, survey reports and articles related to various aspects of herpetological research in magazines or non-peer reviewed media.
2. Allain, S. J. R. & Goodman, M. J. (2015). Cambridgeshire Amphibian Survey Report 2014. Nature in Cambridgeshire, 57, 101-108.
3. Stark, T., Viernum, S., Allain, S., Spiller, M. & Reeves, V. (2015). Crowdfunding for Chytrid 2.0 (Batrachochytrium salamandrivorans) in Belgium. Froglog, 116, 20.
4. Allain, S. J. R. & Goodman, M. J. (2016). Lamination as a method of preserving reptile sloughs. Herpetological Bulletin, 138, 29-30.
5. Allain, S. J. R. & Smith, L. T. (2016). Newt mortalities on an urban cycle path. Herpetological Bulletin, 138, 27-28.
6. Allain, S. J. R. & Goodman, M. J. (2017). Cambridgeshire Amphibian Survey Report 2015. Nature in Cambridgeshire, 59, 35-44.
7. Allain, S. J. R., Smith, L. T. & Miller, G. J. (2017). Introduced non-native amphibians and reptiles in Cambridgeshire 2010-2016. Nature in Cambridgeshire, 59, 44-50.
8. Allain, S., Stark, T. & Viernum, S. (2017). An Update on Crowdfunding for Chytrid 2.0. FrogLog, 118, 29.
9. Allain, S. J. R. & Goodman, M. J. (2017). A case of xanthochromism in the common frog (Rana temporaria). Herpetological Bulletin, 139, 39-40.
10. Allain, S. J. R. & Goodman, M. J. (2017). Using call playbacks to investigate a population of non-native midwife toads Alytes obstetricans (Laurenti, 1768) in Cambridge, UK. Herpetological Bulletin, 140, 28-30.
11. Allain, S. J. R. & Smith, L. T. (2017). New records of paedomorphic smooth newts (Lissotriton vulgaris) at a site in Cambridgeshire, U.K. Herpetological Bulletin, 141, 40.
12. Allain, S. J. R. & Goodman, M. J. (2017). Monitoring a Non-native Amphibian in Cambridge, UK. FrogLog, 119, 36-37.
13. Allain, S. J. R. (2017). The use of roadside gabions as refugia by reptiles in Kota Kinabalu, Sabah, Malaysia. Captive & Field Herpetology, 1(1), 7-9.
14. Allain, S. J. R. & Goodman, M. J. (2017). Absence of chytrid fungus (Batrachochytrium dendrobatidis) in an introduced population of the common midwife toad (Alytes obstetricans) in Cambridge, UK. Herpetological Bulletin, 142, 40-41.
15. Allain, S. J. R. (2018). Comments on the Swabbing of Anuran Tadpoles for the Detection of Infectious Diseases. FrogLog, 120, 40.
16. Allain, S. J. R. & Goodman, M. J. (2018). The absence of the amphibian chytrid fungi in the common midwife toad (Alytes obstetricans) from an introduced population in Cambridge, UK. Herpetology Notes, 11, 451-454.
17. Allain, S. J. R. & Goodman, M. J. (2018). Cambridge Amphibian Survey Report 2016. Nature in Cambridgeshire, 60, 31-36.
18. Allain, S. J. R. & Goodman, M. J. (2018). New records of Palmate Newts (Lissotriton helveticus) in Cambridge. Nature in Cambridgeshire, 60, 36-37.
19. Allain, S. J. R. & Goodman, M. J. (2018). Digital measuring spoons, another instrument in the field herpetologist’s tool-box? Herpetological Bulletin, 144, 26-27.
20. Cunningham, O. K. E. & Allain, S. J. R. (2018). A Description of the Successful Hatching of the Fijian Banded Iguana (Brachylophus bulabula) and Observations on Vermiculite Moisture. Captive & Field Herpetology, 2(1), 12-15.
21. Allain, S. J. R. (2018). In the shadow of mammals: the use of camera traps to investigate the basking behaviour of Agama mwanzae in the Serengeti National Park, Tanzania. Herpetological Bulletin, 145, 19-21.
22. Allain, S. J. R. & Bateman, T. C. B. (2018). Ixodid Ticks on Oertzen’s Rock Lizard (Anatololacerta oertzeni) on Ikaria, Greece, with Notes on the Island’s Reptiles. Reptiles & Amphibians, 25(3), 176-179.
23. Allain, S. J. R. & Goodman, M. J. (2018). The Defense Mechanism of Kuhl’s Stream Frog (Limnonectes cf. kuhlii). Reptiles & Amphibians, 25(3), 180-181.
24. Allain, S. J. R. (2019). Now you see me, now you don't. Frontiers in Ecology & the Environment, 17(1), 31.
25. Saucedo, B., Garner, T. W. J., Kruithof, N., Allain, S. J. R., Goodman, M. J., Cranfield, R. J., Sergeant, C., Vergara, D. A., Kik, M. J. L., Forzán, M. J., van Beurden, S. J. & Gröne, A. (2019). Common midwife toad ranaviruses replicate first in the oral cavity of smooth newts (Lissotriton vulgaris) and show distinct strain-associated pathogenicity. Scientific Reports, 9, 4453.
26. Allain, S. J. R., Gandola, R. & Wilkinson, J. W. (2019). One or multiple origins of midwife toads in the UK? FrogLog, 121, 23.
27. Allain, S. J. R. (2019). Comments on the collection of buccal swabs from amphibians. FrogLog, 121, 29.
28. Allain, S. J. R., Goodman, M. J. & Jopling, A. D. (2019). Notes on the successful mitigation-driven translocation of slow worms (Anguis fragilis) at Wandlebury Country Park. Nature in Cambridgeshire, 61, 67-69.
29. Allain, S. J. R. & Goodman, M. J. (2019). New records of midwife toads (Alytes obstetricans) in Cambridgeshire. Nature in Cambridgeshire, 61, 69-70.
30. Allain, S. J. R. & Goodman, M. J. (2019). Cambridge Amphibian Survey Report 2017. Nature in Cambridgeshire, 61, 70-75.
31. Cunningham, O. K. E. & Allain, S. J. R. (2019). First record of intraspecific oophagy in the common leaf-tailed gecko Uroplatus fimbriatus (SCHNEIDER, 1797). Captive & Field Herpetology, 3(1), 19-21.
32. Allain, S. J. R. (2019). Mining Flickr: a method for expanding the known distribution of invasive species. Herpetological Bulletin, 148, 11-14.
33. Allain, S. J. R. & Smith, L. T. (2019). A suspected case of infection by Amphibiocystidium sp. at a site in Cambridgeshire, UK. SAURIA, 41(3), 58–60.
34. Allain, S. J. R. & Mattea, R. G. (2019). An observation of extreme necrosis in the Italian wall lizard (Podarcis sicula). Reptiles & Amphibians, 26(2), 102.
35. Bray, I. H. & Allain, S. J. R. (2019). Are squash boxes necessary? The use of dorsal photographs to determine snout-vent lengths of slow worms (Anguis fragilis). Reptiles & Amphibians, 26(2), 165-166.
36. Allain, S. J. R. & Duffus, A. L. J. (2019). Emerging Infectious Disease Threats to European Herpetofauna. Herpetological Journal. 29(4), 189-206.
37. Oberhofer, S. J. & Allain, S. J. R. (2019). Autumn activity in an introduced population of Podarcis muralis ssp. in Liechtenstein. L@CERTIDAE, 2019(7), 54–57.
38. Smales, L. R., Allain, S. J. R., Wilkinson, J. W. & Harris, E. (2020). A new species of Pseudoacanthocephalus (Acanthocephala: Echinorhynchidae) from the guttural toad, Sclerophrys gutturalis (Bufonidae), introduced into Mauritius, with comments on the implications of the introductions of toads and their parasites into the United Kingdom. Journal of Helminthology, 94, e119.
39. Allain, S. J. R. & Goodman, M. J. (2020). Notes on the Herpetofauna of the Singapore Botanic Gardens. Reptiles & Amphibians, 26(3), 216–218.
40. Allain, S. J. R. & Goodman, M. J. (2020). An Incidence of Road Mortality in the Asian Water Monitor (Varanus salvator) in Kota Kinabalu, Sabah, Malaysia. Reptiles & Amphibians, 26(3), 219–220.
41. Grassby-Lewis, R. J. & Allain, S. J. R. (2020). Observation of Arboreal Behavior in the Malayan Pitviper (Calloselasma rhodostoma). Reptiles & Amphibians, 27(1), 79–80.
42. Mattea, R. G. & Allain, S. J. R. (2020). An Observation of Nocturnal Activity in a Central-Italian Asp (Vipera aspis francisciredi). Reptiles & Amphibians, 27(1), 87–88.
43. Clemens, D. J. & Allain, S. J. R. (2020). New records of Aesculapian snakes (Zamenis longissimus) in South Wales, UK. Herpetological Bulletin, 152, 30-31.
44. Clemens, D. J. & Allain, S. J. R. (2020). First evidence of Fogo skink (Chioninia fogoensis) introduction to Madeira. Herpetological Bulletin, 152, 40-41.
45. Harnkness, H. & Allain, S. J. R. (2020). A case of leucism in the slow worm (Anguis fragilis). Herpetological Bulletin, 152, 43.
46. Thomas, O., Allain, S. J. R. & Sagar, P. (2020). Predation and ingestion of a viviparous lizard (Zootoca vivipara) by the common wall lizard (Podarcis muralis) in Dorset, UK. Herpetological Bulletin, 152, 44.
47. West, P. & Allain, S. J. R. (2020). A case of erythrism in the common frog (Rana temporaria). Reptiles & Amphibians, 27(2), 331–332.
48. Shin, Y., Yikweon, J., Allain, S. J. R. & Borzée, A. (2020). Catalogue of herpetological specimens of the Ewha Womans University Natural History Museum (EWNHM), Republic of Korea. ZooKeys, 965, 103–139.
49. Allain, S. J. R. (2020) Snakes in the grass: the misidentification of adders in Cambridgeshire. Nature in Cambridgeshire, 62, 51-54.
50. Allain, S. J. R. & Bray, I. H. (2020). The ongoing monitoring of slow worms (Anguis fragilis) at Wandlebury Country Park. Nature in Cambridgeshire, 62, 54-56.
51. Allain, S. J. R. & Garcia, G. (2020). Promoting research, education and conservation: 9th World Congress of Herpetology. Herpetological Bulletin, 153, 1-2.
52. Mattea, R. G. & Allain, S. J. R. (2020). CHELONOIDIS DENTICULATUS (Yellow-footed Tortoise). DIET. Herpetological Review, 51(3), 576.
53. Mattea, R. G. & Allain, S. J. R., Culebras, J. & Barahona-Velastegui, A. M. (2020). HYLOSCIRTUS MASHPI (Mashpi Stream Treefrog). Herpetological Review, 51(4), 769-770.
54. Allain, S. J. R. & Clemens, D. J. (2020). Online news reports as a source of data on the occurrence and frequency of stowaway reptiles and amphibians entering Great Britain. Herpetological Bulletin, 154, 8-11.
55. Lewarne, B. & Allain, S. J. R. (2020). The unnatural history of cave olms Proteus anguinus in England. Herpetological Bulletin, 154, 18-19.
56. Baker, J. M. R. & Allain, S. J. R. (2020). Photographic Identification of Individual Barred Grass Snakes Natrix helvetica. Transactions of the Norfolk & Norwich Naturalists’ Society, 53(1), 67-70.
57. Allain, S. J. R., Gandola, R., Tighe, A. & Wilkinson, J. W. (2021). An investigation into the provenance of Bedford’s midwife toads. Bedfordshire Naturalist, 74(1), 48-52.
58. Allain, S. & Duffus, A. (2021). Special Issue on Emerging Diseases and Infections in Herpetofauna. FrogLog, 122, 14.
59. Allain, S. J. R., West, P. & Griffiths, S. (2021). Observations of polymely in the smooth newt (Lissotriton vulgaris) and the palmate newt (L. helveticus). Herpetological Bulletin, 155, 21-22.
60. Thomas, O., Allain, S. J. R. & Owen, H. (2021). Predation and ingestion of a juvenile barred grass snake (Natrix helvetica) by a rainbow trout (Oncorhynchus mykiss) in Lot, France. Herpetological Bulletin, 156, 41-42.
61. Clemens, D. J. & Allain, S. J. R. (2021). An unusually high number of Italian wall lizards Podarcis siculus campestris entering Great Britain as stowaways. Herpetological Bulletin, 156, 43.
62. Allain, S. J. R., Goodman, M. J. & Wilkinson. J. W. W. (2021). A series of unlikely events: from washing machine to new species. Herpetological Bulletin, 156, 45-46.
63. Thomas, O. & Allain, S. J. R. (2021). A Review of Prey Taken by Anacondas (Squamata: Boidae: Eunectes). Reptiles & Amphibians, 28(2), 329-334.
64. Allain, S. J. R. & Shimbov, M. I. (2021). The mystery of the Thorney midwife toads (Alytes obstetricans). Nature in Cambridgeshire, 63, 26-28.
65. Shimbov, M. I. & Allain, S. J. R. (2021). Male common midwife toad Alytes obstetricans depositing eggs in a flowerpot saucer in a suburban garden? Herpetological Bulletin, 157, 38-39.
66. Clemens, D. J., Rose, L. G. & Allain, S. J. R. (2021). Tadpoles of the midwife toad Alytes obstetricans scavenging carrion. Herpetological Bulletin, 157, 46.
67. Allain, S. J. R. & Lynn, V. J. (2021). Distribution of the alpine newt Ichthyosaura alpestris in Great Britain updated using social media. Herpetological Bulletin, 158, 28-31.
68. Duffus, A. L. J., Hughes, D. F., Kautz, A., Allain, S. J. R. & Meshaka, W. E. (2022). Repeated Sampling Of Wild Individuals Reveals Ophidiomyces ophidiicola Infection Dynamics In A Pennsylvania Snake Assemblage. Journal of Wildlife Diseases, 58(2), 290-297.
69. Lynn, V. J. & Allain, S. J. R. (2022). Observations on the climbing behaviour of the smooth newt Lissotriton vulgaris and great crested newt Triturus cristatus in south-east England. Herpetological Bulletin, 160, 25-26.
70. Goodman, M. J., Clemens, D. J., Rose, L. G. & Allain, S. J. R. (2022). Limb malformations in introduced populations of midwife toad Alytes obstetricans in Great Britain. Herpetological Bulletin, 161, 31-33.
71. Shimbov, M. I. & Allain, S. J. R. (2022). Smooth newt Lissotriton vulgaris preyed upon by European starling Sturnus vulgaris. Herpetological Bulletin, 161, 46.
72. Allain, S. J. R., Duffus, A. L. J. & Marshang, R. E. (2022). Editorial: Emerging infections and diseases of herpetofauna. Frontiers in Veterinary Science, 9, 909616.
73. Squire, B. & Allain, S. J. R. (2022). BUFO BUFO (Common Toad). UNUSUAL SPAWN. Herpetological Review, 53(3), 465-466.
74. Allain, S. J. R. & Lynn, V. J. (2023). Records of terrapins (Emydidae & Kinosternidae: Testudines) in Tunbridge Wells, Kent. Transactions of the Kent Field Club, 21, 49-55.
75. Allain, S. J. R., Clemens, D. J. & Thomas, O. (2023). Taste the Rainbow: A Review of Color Abnormalities Affecting the Herpetofauna of the British Isles Reptiles & Amphibians, 30, e18470.
76. Davies, N. & Allain, S. J. R. (2023). Smooth newts Lissotriton vulgaris as more than just occasional items in the diet of the Eurasian kingfisher Alcedo atthis. Herpetological Bulletin, 164, 43.
77. Robbemont, J., van Veldhuijzen, S., Allain, S. J. R., Ambu, J., Boyle, R., D., Cvijanović, M., Dufresnes., C., Ennis, C., Gandola, R., Jablonski., D., Julian, A., Kranželić, D., Lukanov, S., Martínez-Solano, I., Montgomery, R., Naumov, B., O’Neill, M., North, A., Pabijan, M., Puschendorf., R., Salvi, D., Schmidt, B., Sotiropoulos, K., Stanescu, F., Stanković, D., Stapelton, S., Šunje, E., Szabolcs, M., Vacheva, E., Willis, D., Zimić, A., France, J., Meilink, W. R. M., Stark, T., Struij, R. P. J. H., Theodoropoulos, A., de Visser, M. C. & Wielstra, B. (2023). An extended mtDNA phylogeography for the alpine newt illuminates the provenance of introduced populations. Amphibia-Reptilia, 44(3), 347-361.
78. Goodman, M. J., Clemens, D. J., Shimbov, M. I. & Allain, S. J. R. (2023). No amphibian chytrid fungus Batrachochytrium dendrobatidis detected in four introduced populations of the midwife toad Alytes obstetricans in eastern England. Herpetological Bulletin, 165, 1-4.
79. Lewarne, B. & Allain, S. J. R. (2023). Attempted predation of a cave olm Proteus anguinus by a dice water snake Natrix tessellata, in Bosnia and Herzegovina. Herpetological Bulletin, 165, 34-35.
80. Lymer, M. & Allain, S. J. R. (2024). The barred grass snake Natrix helvetica as a potential predator of sand martin Riparia riparia chicks. Herpetological Bulletin, 167, 38-39.
81. Allain, S. J. R., Leech, D. I., Hopkins, K., Seilern-Moy, K., Fernandez, J, R.-R., Griffiths, R. A. & Lawson, B. (2024). Characterisation, prevalence and severity of skin lesions caused by ophidiomycosis in a population of wild snakes. Scientific Reports, 14, 5162.
82. Allain, S. J. R. & Deaton, L. J. (2024). The Common Midwife Toad Alytes obstetricans in Norfolk. Transactions of the Norfolk & Norwich Naturalists’ Society.
83. Allain, S. J. R. & Phillips, N. (2024). Observations of a neotenous population of Smooth Newts Lissotriton vulgaris from Norfolk. Transactions of the Norfolk & Norwich Naturalists’ Society.
84. Goodman, M. J., Shimbov, M. I. & Allain, S. J. R. (2024). A third population of the common midwife toad (Alytes obstetricans) present within Cambridgeshire. Nature in Cambridgeshire, 66.
85. Shimbov, M. I., Goodman, M. J. & Allain, S. J. R. (2024). First records of the non-native alpine newt (Ichthyosaura alpestris) in Cambridgeshire. Nature in Cambridgeshire, 66.
1. Allain, S. J. R. (2015). 'The use of bottle traps at a colony of great crested newts (Triturus cristatus) at a site in Cambridgeshire to investigate aquatic microhabitat preferences', Bachelor's Dissertation, Anglia Ruskin University. 2. Allain, S. J. R. (2017). 'An expanded geographical range of the amphibian chytrid fungus (Batrachochytrium dendrobatidis) in Madagascar', Master's Thesis, Imperial College London. 3. Allain, S. J. R. (2018). 'Investigating the conservation status and changes in species temporal distributions of Asian amphibians using museum specimens', Master's Thesis, Imperial College London. 4. Allain, S. J. R. (2022). 'Population dynamics, disease and other drivers in a wild snake population', PhD Thesis, University of Kent.
1. Allain, S. J. R. (2024). Book Review: Tadpole Hunter. Reptiles & Amphibians, 31(1), e21657.
1. Allain, S. J. R. (2021). The Challenges of Reptile Conservation in Great Britain. Indigo Magazine, 13, 28-31.
2. Allain, S. J. R. (2022). A quick hop through frog stamps of the world. Stamp Collector, 4(4), 60-63.
3. Allain, S. J. R. (2022). Big fangs coming in small packages. Stamp Magazine, 88(7), 50-54.
1. Allain, S. J. R. (2018). The making of an amphibian conservation biologist. BES Bulletin, 49(1), 64-65.
2. Price, J. (2018). Working for nature: Steven Allain - Amphibian and reptile surveys, Cambridgeshire. BBC Wildlife, 36(11), 115.
